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2002-2003 General Catalog
University of California, Riverside

ENTOMOLOGY

Subject abbreviation: ENTM


Timothy D. Paine, Ph.D., Chair
J. Daniel Hare, Ph.D., Vice Chair
Department Office, 175 Entomology
 insects.ucr.edu
Graduate Student Affairs (800) 735-0717 or (909) 787-5621
 insects.ucr.edu/programs/graduate.html
Undergraduate Advisor (909) 787-4562
 insects.ucr.edu/programs/undergraduate.html

Faculty E-mails

Professors
Michael E. Adams, Ph.D. (Entomology/Cell Biology and Neuroscience)
Nancy E. Beckage, Ph.D. (Entomology/Cell Biology and Neuroscience)
Thomas S. Bellows, Jr., Ph.D.
Ring T. Cardé, Ph.D. Alfred M. Boyce Chair in Entomology
Brian A. Federici, Ph.D.
J. Daniel Hare, Ph.D.
Robert F. Luck, Ph.D.
Jocelyn G. Millar, Ph.D.
Thomas A. Miller, Ph.D.
Joseph G. Morse, Ph.D.
Mir S. Mulla, Ph.D.
Bradley A. Mullens, Ph.D.
Timothy D. Paine, Ph.D.
Thomas M. Perring, Ph.D.
John D. Pinto, Ph.D.
Alexander Raikhel, Ph.D.
Richard A. Redak, Ph.D.
Michael K. Rust, Ph.D.
S. Nelson Thompson, Ph.D.
John T. Trumble, Ph.D.
Professors Emeriti
Martin M. Barnes, Ph.D.
Leland R. Brown, Ph.D.
Glenn E. Carman, Ph.D.
Richard D. Goeden, Ph.D.
E. Fred Legner, Ph.D.
Ralph B. March, Ph.D.
James A. McMurtry, Ph.D.
Earl R. Oatman, Ph.D.
Louis A. Riehl, Ph.D.
Vernon M. Stern, Ph.D.
Associate Professors
Peter W. Atkinson, Ph.D.
John M. Heraty, Ph.D.
Richard Stouthamer, Ph.D.
P. Kirk Visscher, Ph.D.
Gregory P. Walker, Ph.D.
William E. Walton, Ph.D.
••
Lecturer
Daniel González, Ph.D.
Heather Costa, Ph.D. Ornamental Crops
Elizabeth Grafton-Cardwell, Ph.D. Pest Management
Mark Hoddle, Ph.D. Biological Control
John H. Klotz, Ph.D. Urban Entomology
Robert Krieger, Ph.D. Toxicology
Nick Toscano, Ph.D. Pest Management

MAJOR

The Department of Entomology offers undergraduate programs leading to either the B.S. or the B.A. degree. The B.S. degree offers students with a strong interest in the natural sciences an opportunity to emphasize this aspect of their education. The B.A. degree is available to students who wish to obtain a broader background in the humanities and social sciences than is required of students in the B.S. program.

Counseling and information on the program and course requirements is provided by the departmental Undergraduate Advisor, Dr. Thomas M. Perring, 225 Entomology.

Degree Requirements

University Requirements

See the Undergraduate Studies section for requirements that all students must satisfy.

College Requirements

See Degree Requirements, College of Natural and Agricultural Sciences, in the Undergraduate Studies Section, for requirements that students must satisfy.

Some of the following requirements for the major may also fulfill some of the college's breadth requirements. Consult with a department advisor for course planning.

Major Requirements

The major requirements for both the B.A. and the B.S. degrees in Entomology are as follows:

1. Lower-division requirements (47 units)

    a) BIOL 005A, BIOL 05LA, BIOL 005B, BIOL 005C
    b) PHYS 002A, PHYS 002B, PHYS 002C, PHYS 02LA, PHYS 02LB, PHYS 02LC
    c) MATH 009A, MATH 009B
    d) CHEM 001A, CHEM 001B, CHEM 001C
2. Upper-division requirements (62 units)
    a) ENTM 100/BIOL 100, ENTM 109, ENTM 112/BIOL 112/BPSC 112, ENTM 127/BIOL 127, ENTM 173/BIOL 173
    b) Eighteen (18) additional units of entomology electives, which may include up to 4 units of ENTM 190, ENTM 197, or ENTM 199H
    c) BCH 100
    d) BIOL 102
    e) BIOL 104/BPSC 104
    f) CHEM 112A, CHEM 112B, CHEM 112C
3. Statistics (2 or 5 units): STAT 020 or STAT 100A

BIOL 151 and BIOL 175 are suggested in order to acquire a background in the life sciences appropriate for an Entomology major.

For students intending to specialize at the graduate level in insect toxicology or insect physiology, biochemistry, and molecular biology, it is recommended that the BCH 110A, BCH 110B, and BCH 110C sequence and BCH 102 be substituted in place of an equal number of upper-division course units in life sciences. Due to course content overlap, credit is not awarded for BCH 110A, BCH 110B, or BCH 110C if it has already been awarded for BCH 100.

Entomology Sample Program

The Department of Entomology also participates in the Biological Sciences major. See the Biological Sciences listing in this catalog for information about its Entomology track.

Minor

The Department of Entomology offers a minor in Entomology designed to allow the student the freedom to pursue areas of particular interest.

The minor consists of no less than 20 and no more than 28 units of Entomology courses to be selected as follows:

  1. ENTM 100/BIOL 100
  2. Select from the following upper-division Entomology courses to complete unit requirement: ENTM 109, ENTM 112/BIOL 112/BPSC 112, ENTM 114, ENTM 124, ENTM 126, ENTM 126L, ENTM 127/BIOL 127, ENTM 128, ENTM 129, ENTM 129L, ENTM 132, ENTM 133, ENTM 162/ BIOL 162, ENTM 173/BIOL 173, ENTM 190, ENTM 197, ENTM 199H
  3. No more than 4 units of ENTM 190, ENTM 197, or ENTM 199H, either solely or in combination, may be applied toward the unit requirement.
  4. Students may count no more than 8 units toward both their major and minor fields of study.

See Minors under the College of Natural and Agricultural Sciences in the Undergraduate Studies section of this catalog for additional information on minors.

GRADUATE PROGRAM

The Department of Entomology offers programs leading to the M.S. (thesis plan) and Ph.D. degrees with specialization in, but not restricted to, the following areas of study:

• Arthropod vectors of plant pathogens
• Behavior
• Biochemistry and physiology
• Biological control
• Chemical ecology
• Ecology and evolution
• Integrated pest management
• Insect–plant interactions
• Medical and veterinary entomology
• Molecular entomology
• Nematology
• Neuroscience
• Pathology
• Pesticide toxicology
• Systematics
• Urban entomology

Information on participating faculty and their research specializations may be found at insects.ucr.edu and in the brochure Graduate Studies in Entomology. The Supplementary Information Pamphlet provides detailed information for completing the graduate program. These publications may be obtained from the Biological Sciences Graduate Student Affairs Center, 1151 Batchelor Hall, (800) 735-0717. University requirements for the M.S. and Ph.D. degrees are given in the Graduate Studies section of this catalog.

Admission Students must have a bachelor's degree with a major in Entomology, a biological science, Chemistry, Biochemistry, or a suitable equivalent. Course work is required in inorganic and organic chemistry, including laboratories, and general physics.

Additional admission requirements for M.S. and Ph.D. students consist of the equivalent of 30 quarter units of life sciences other than entomology, including one course in general biology and genetics. A course in biochemistry may serve as an elective in the life sciences. Students specializing in insect biochemistry, insect physiology, molecular entomology, neuroscience, or toxicology may substitute courses in organic, physical, and biological chemistry; toxicology; and pharmacology for courses in life sciences except for the equivalent of a one-year introductory course in general biology.

The department requires GRE General Test scores (verbal, quantitative, and analytical) and scores from one of the advanced tests. This requirement applies only to U.S. citizens and to international applicants residing in the United States at the time of application. The department strongly recommends that international students residing outside of the United States at the time of application also submit GRE scores.

All applicants whose first language is not English are required to complete the Test of English as a Foreign Language (TOEFL). A minimum score of 550 is required on the paper-based exam. A minimum score for the computer-based exam is 213.

Opportunities for Interdisciplinary Graduate Study

Faculty from the Department of Entomology also participate in unique graduate specializations in Cell, Molecular, and Developmental Biology, Cell Biology and Neuroscience, Environmental Toxicology, Evolution and Ecology, and Genetics, which draw on the strengths of distinguished scientists from several units. For further information concerning work in these areas, see the respective program descriptions in the Curricula and Courses section of this catalog or contact the Biological Sciences Graduate Student Affairs Center, at (800) 735-0717.

Normative Time to M.S. 6 quarters

Normative Time to Ph.D. 17 quarters


LOWER-DIVISION COURSES

ENTM 010. Natural History of Insects. (4) F,W,S Lecture, three hours; demonstrations, one hour. A study of the fascinating world of insects and of their impact on man; designed for non-entomology majors. Living and preserved insects and many other visual aids are used. Federici, Gonzalez, Luck, Morse, Mullens, Paine, Perring, Redak, Rust, Trumble

ENTM 020. Bees and Beekeeping. (4) F, Odd Years Lecture, three hours; discussion, one hour. Fundamentals of keeping honey bees, their fascinating social behavior, and their economic importance as pollinators of agricultural crops and as producers of honey and other products. Demonstrations of bee biology and behavior, with colonies of bees, and of beekeeping techniques, equipment, and extraction of honey. Visscher


UPPER-DIVISION COURSES

ENTM 100. General Entomology. (4) F Lecture, three hours; laboratory, three hours. Prerequisite(s): BIOL 005B, BIOL 005C, or equivalents; or consent of instructor. Introductory study of insects, Earth's most diverse group of animals (75 percent of animal species are insects). Lecture covers the anatomy, physiology, ecology, behavior, and diversity of insects. Laboratory focuses on insect identification. Cross-listed with BIOL 100. Walker

ENTM 109. Field Entomology. (4) S Laboratory, four hours; field, eight hours. Prerequisite(s): BIOL 100/ENTM 100 or equivalents or consent of instructor. Study and field collection of insects in selected ecological communities from the diversity of life zones comprising Southern California. Students prepare specimens collected to professional standards, identify specimens, and submit their collections for grading and incorporation into the Department of Entomology's teaching and research collections. Stouthamer

ENTM 112. Systematics. (4) F Lecture, three hours; discussion, one hour. Prerequisite(s): BIOL 005C or equivalent. Principles and philosophy of classification: phylogenetic and phenetic methods, species concepts, taxonomic characters, evolution, hierarchy of categories, and nomenclature. Cross-listed with BIOL 112 and BPSC 112. Heraty, Kim

ENTM 114. Aquatic Insects. (4) S, Even Years Lecture, three hours; laboratory, three hours. Prerequisite(s): BIOL 005A, BIOL 005B, BIOL 005C; or consent of instructor. Investigates aquatic insects as nutrient cyclers, pollution indicators, disease vectors, and fish food. Involves identification of major orders and families, morphological and physiological adaptations, and life history strategies. Laboratory emphasizes identification (collection) and includes a group field ecology project and two weekend field trips. Mullens, Walton

ENTM 124. Agricultural Entomology. (4) F, Odd Years Laboratory, four hours; field, eight hours. Prerequisite(s): BIOL 100/ENTM 100 or equivalent or consent of instructor. Identification, life history, ecology, distribution, and management of key pest and beneficial species learned through field observation, discussions with industry representatives, and laboratory study. Detailed notes and collections from field trips to all major growing regions of Southern California form the basis for laboratory discussion. Perring

ENTM 126. Medical and Veterinary Entomology. (3) W, Odd Years Lecture, three hours. Prerequisite(s): BIOL 005A, BIOL 005B, BIOL 005C; or consent of instructor. Biology, ecology, and management of arthropods affecting human and animal health. Arthropods as direct pests and vectors of important diseases (e.g., malaria, plague). Disease epidemiology and prevention and control of pests and associated diseases are discussed. Mulla, Mullens

ENTM 126L. Laboratory in Medical and Veterinary Entomology. (2) W, Odd Years Laboratory, six hours. Prerequisite(s): BIOL 005A, BIOL 005B, BIOL 005C; or consent of instructor. Concurrent or previous enrollment in ENTM 126 is recommended. Identification of arthropods affecting humans and animals. Practical epidemiological exercises, including age-grading, blood meal and pathogen identification in vectors, vector capacity assessment, bioassay procedures, and sampling. Field trip(s) to animal production and mosquito abatement and research facilities are scheduled. Mulla, Mullens

ENTM 127. Insect Ecology. (4) Lecture, three hours; discussion, one hour. Prerequisite(s): BIOL 005A, BIOL 05LA, BIOL 005B, BIOL 005C, CHEM 001C or CHEM 01HC, CHEM 112C, MATH 009B or MATH 09HB, PHYS 002C, PHYS 02LC, BCH 100 or BCH 110A, one course in statistics; or consent of instructor. Principles of insect ecology. Topics, with examples emphasizing the Arthropoda, include factors governing population growth; ecological and evolutionary interactions with hosts, competitors, and natural enemies; structure of ecological communities; and adaptations to different environments. Cross-listed with BIOL 127. Credit is not allowed for both BIOL 117 and BIOL 127/ENTM 127. Bellows, Walton

ENTM 128. Chemistry and Toxicology of Insecticides. (3) F, Even Years Lecture, three hours. Prerequisite(s): a course in organic chemistry, BIOL 100/ ENTM 100; or consent of instructor. Chemical properties and reactions of insecticides and acaricides and their modes of action and biochemical behavior in animal and plant systems. Miller

ENTM 129. Introduction to Biological Control. (2) F Lecture, two hours. Prerequisite(s): BIOL 100/ ENTM 100 or consent of instructor. Principles and methods of biological control; biology and behavior of entomophagous insects; historical review and critique of important world projects. Stouthamer

ENTM 129L. Introduction to Biological Control Laboratory. (2) F Laboratory, six hours. Prerequisite(s): ENTM 129 (it is strongly recommended that ENTM 129L be taken concurrently with ENTM 129). Laboratory identification of entomophagous insects; experiments designed to illustrate various types of parasitism; familiarization with mass rearing and culture techniques for entomophagous insects. Heraty

ENTM 132. Taxonomy of Immature Insects. (4) S, Alternate Years Lecture, two hours; laboratory, six hours. Prerequisite(s): BIOL 100/ENTM 100 or consent of instructor. Morphology and identification of the immature stages of insects. Pinto

ENTM 133. Urban Entomology. (4) S, Even Years Lecture, three hours; laboratory, three hours. Prerequisite(s): BIOL 100/ENTM 100 or consent of instructor. Biology and management of arthropod pests of the urban- industrial community with an emphasis on structural, household, and stored product pests. Exercises on the recognition and identification of these pests, their life histories, and strategies for their control. Rust

ENTM 162. Insect Behavior. (4) F Lecture, four hours. Prerequisite(s): BIOL 100 /ENTM 100; or BIOL 005A, BIOL 005B, and BIOL 005C; or consent of instructor. An analysis of the mechanisms that cause and control behavioral reactions of insects. Emphasis on ethological and physiological knowledge concerning orientation mechanisms, communication systems, learning, and the role of the nervous system in integrating behavior in insects. Cross-listed with BIOL 162. Cardé, Visscher

ENTM 173. Insect Physiology. (4) Lecture, three hours; laboratory, three hours. Prerequisite(s): BIOL 005A and BIOL 005B or equivalents; CHEM 112A, CHEM 112B, CHEM 112C or equivalents; or consent of instructor. Introduction to principles of insect physiology. Subjects include growth, development and hormones, cuticle, nervous system, circulation, respiration, digestion, nutrition, excretion, reproduction, water balance, and temperature relations. Prior knowledge of insects is not assumed. Cross-listed with BIOL 173. Miller, Thompson

ENTM 190. Special Studies. (1-5) F,W,S Prerequisite(s): consent of instructor. Directed studies in specialized fields in entomology such as insects affecting subtropical fruits, deciduous fruits and nuts, floricultural crops and turf, vegetable and field crops, forest and ornamental trees and shrubs, stored products, and households. Course is repeatable.

ENTM 197. Research for Undergraduates. (1-4) F,W,S Prerequisite(s): consent of instructor. Directed original research and preparation of written report. Course is repeatable.

ENTM 199H. Senior Honors Research. (1-5) F,W,S Laboratory, three to fifteen hours. Prerequisite(s): senior status and consent of instructor; a GPA of 3.5 or better in entomology courses and 3.2 in all University course work. Research in entomology under supervision of a faculty member in entomology. The student will submit a written report. Course is repeatable.


GRADUATE COURSES

ENTM 204. Advanced Insect Ecology. (3) F Lecture, three hours. Prerequisite(s): an upper-division course in population and community ecology or consent of instructor. Graduate-level introduction to principles of insect ecology, genetics, evolution, and behavior. Topics include insect population dynamics and community interactions, behavioral interactions of insects with their environment, and genetics of geographic variation and adaptation of insect populations. Hare, Millar, Visscher

ENTM 205. Insect Morphology. (3) W Lecture, three hours. Prerequisite(s): BIOL 100/ENTM 100 or an equivalent upper-division general entomology course or consent of instructor. Principles of insect morphology, with emphasis on functional systems and morphological characters of phylogenetic importance and adaptive significance to insects; comparative anatomy of extinct and living insect groups; insect phylogenetic relationships. Pinto

ENTM 205L. Insect Taxonomy Laboratory. (2) W Laboratory, six hours. Prerequisite(s): BIOL 100/ENTM 100 or an equivalent upper-division general entomology course, and concurrent enrollment in ENTM 205, or consent of instructor. Insect taxonomy, stressing the characteristics of the major taxa and identification to the level of family. Pinto

ENTM 206. Insect Physiology and Biochemistry. (3) S Lecture, three hours. Prerequisite(s): upper-division courses in general entomology and general biochemistry or consent of instructor. Graduate-level introduction to the physiology and biochemistry of insect systems. Topics covered include basics of growth and development, reproduction, digestion, nutrition, metabolism, respiration, circulation, ion and water balance, nervous and muscular systems, circadian rhythms. Adams, Thompson

ENTM 206L. Insect Physiology Laboratory. (2) S Laboratory, six hours. Prerequisite(s): upper-division courses in general entomology and general biochemistry or consent of instructor. Laboratory experiments and demonstrations designed to illustrate basic principles of insect physiology and to provide opportunities for students to acquire basic methodological skills in chromatography, pharmacology, bioassay, and use of radiotracers. Adams, Millar

ENTM 207. Arthropod Vectors in Relation to Plant Disease. (4) S, Even Years Lecture, two hours; laboratory, six hours. Prerequisite(s): BIOL 100/ENTM 100, BIOL 120/MCBL 120/PLPA 120; or consent of instructor. Detailed analyses of interacting mechanisms involved in the transmission of plant pathogens by arthropods. Emphasis on learning through extensive laboratory experimentation. Perring

ENTM 208. Host-Parasite Relationships. (3) Lecture, three hours. Prerequisite(s): BIOL 100/ENTM 100 or BIOL 157 or consent of instructor. Explores the fundamental biochemical and developmental requirements for "successful" host-parasite relationships in insects. Emphasizes wasp and nematode parasites of insects and vector-parasite interactions involved in transmission of parasites in malaria, trypanosoma, and Lyme disease. Cross-listed with BIOL 208. Beckage

ENTM 209. Microtechniques in Insect Morphology. (3) W, Even Years Laboratory, six hours; individual research, three hours. Prerequisite(s): BIOL 005A, BIOL 005B, BIOL 100/ENTM 100; or equivalents; or consent of instructor. Development of research techniques and skills used in the study of insect morphology. Covers the principles of and provides hands-on experience with the following: optical microscopy, scanning electron microscopy, whole-mount slide preparation techniques, morphometric measurement and analysis, scientific illustration, macrophotography, and histological techniques. Walker

ENTM 212. Ecological Systems in Space and Time. (4) Lecture, two hours; discussion, one hour; field, thirty hours per quarter. Prerequisite(s): BIOL 117 or BIOL 152/GEO 152 or equivalent or consent of instructor. Focuses on how ecological systems are interpreted and reconciled at the community, landscape, and paleontological scales and on the role of extrinsic factors operating at each of these scales. Examines the historical development of our understanding of ecological systems at various scales. Cross-listed with BIOL 212 and GEO 212.

ENTM 219. Theory of Systematics. (4) Lecture, two hours; discussion, two hours. Prerequisite(s): BIOL 112/BPSC 112/ENTM 112 or equivalent or consent of instructor. Examines topics developed around a series of classical and recent papers on the principles, philosophy, and methodology of modern systematics and phylogenetic methods. Cross-listed with BIOL 219 and GEO 219. Heraty, Springer

ENTM 224. Advanced Economic Entomology. (4) W Lecture, three hours; discussion, one hour. Prerequisite(s): ENTM 124, BIOL 127/ENTM 127; or consent of instructor. Advanced concepts in insect-pest management and control; ecological bases for control; control by host resistance and by genetic, physical, behavioral, cultural, and chemical means; integrated systems of pest management. Paine

ENTM 226. Insect Development. (3) S, Even Years Lecture, three hours. Prerequisite(s): consent of instructor. Examines the fundamentals of insect development including gametogenesis, fertilization, specification of the body plan, sex determination, larval development, metamorphosis, and formation of the adult, as well as the endocrine and neuroendocrine factors that regulate these processes. Graduate students receive letter grades; undergraduate students receive Satisfactory (S) or No Credit (NC) grades. Beckage

ENTM 227. Insect Population Ecology. (3) W, Odd Years Lecture, three hours. Prerequisite(s): BIOL 127/ENTM 127 or consent of instructor. Recommended: ENTM 129; STAT 100A; STAT 100B or equivalent. Theory of animal population regulation. Factors affecting distribution and abundance of animals with emphasis on examples from the Arthropoda. Luck

ENTM 229. Advanced Biological Control. (4) F, Alternate Years Lecture, three hours; laboratory, three hours. Prerequisite(s): BIOL 127/ENTM 127, ENTM 129, or equivalents, or consent of instructor. The lecture explores theory and practices relating to the use of natural enemies in the suppression of insect, weed, pathogen, and vertebrate populations. The laboratory surveys insect and other natural enemies, their attributes, collection, cultivation, quarantine handling, and field use. Normally letter graded, but students may petition the instructor for a Satisfactory (S) or No Credit (NC) grade. Bellows

ENTM 231. Insect Pathology. (4) S, Even Years Lecture, three hours; laboratory, three hours. Prerequisite(s): BIOL 100/ENTM 100, at least one course in microbiology; or consent of instructor. Consideration of the principles of general insect pathology and microbiology. Detailed study of noninfectious and infectious diseases of insects, diagnosis, epizootiology, physiopathology, symptomatology, and the use of microbial agents in the control of insect pests. Federici

ENTM 232. Molecular Biology of Insects. (3) S, Odd Years Lecture, three hours. Prerequisite(s): BIOL 107A or consent of instructor. Application of molecular biology to entomology and entomological problems. Emphasizes how molecular biological tools are used to understand insect genome organization, pest resistance, transgenic insects, insect behavior, and insect systematics. Atkinson

ENTM 240. Research Methods in Insect Chemical Ecology. (4) W, Odd Years Lecture, three hours; discussion, one hour. Prerequisite(s): BIOL 127/ENTM 127 or ENTM 204 or consent of instructor. Survey of the methods used in the isolation, identification, and bioassay of biologically active natural products. Topics include bioassay design and execution, and microscale chemical separation and identification techniques. Letter grades are assigned to students who present a formal seminar; other students receive Satisfactory (S) or No Credit (NC) grades. Millar

ENTM 241. Insect-Plant Interactions. (4) F, Odd Years Lecture, three hours; discussion, one hour. Prerequisite(s): BIOL 127/ENTM 127 or consent of instructor. Concepts of the development and maintenance of ecological associations between plants and arthropod herbivores in ecological and evolutionary time; organization of arthropod communities on plants; phytochemical basis for the mediation of plant-arthropod associations; coevolution of plants and herbivorous insects; manipulation of plant-arthropod associations in arthropod pest management programs. Hare, Trumble

ENTM 242. Development of Hypotheses and Research Design. (3) Lecture, one hour; discussion, one hour; written work, three hours. Prerequisite(s): graduate standing or consent of instructor. Teaches fundamentals of research topic selection, development of hypotheses, and selection of experimental designs. Students prepare full-length federal grant proposals, then review and rank them in grant panel review format. Millar, Trumble

ENTM 250. Seminar in Entomology. (1) F,W,S Seminar, one hour. A series of lectures by visiting scientists, staff and advanced graduate students on research topics in entomology and allied fields. Graded Satisfactory (S) or No Credit (NC).

ENTM 251. Seminar in Insect-Plant Interactions. (2) W Seminar, two hours. Prerequisite(s): ENTM 241 or consent of instructor. Rigorous examinations and interpretation of recent publications in the area of insect-plant interactions. Subject matter varies from year to year, and the course may be taken more than once for credit. Paine, Trumble, Walker

ENTM 252. Seminar in Insect Behavior. (2) S Seminar, two hours. Prerequisite(s): BIOL 162/ENTM 162 or consent of instructor. An analysis and interpretation of published experimental data dealing with insect behavior, and an attempt to derive general principles underlying behavior. Subject matter varies from year to year, and the course may be taken more than one year for credit. Cardé, Millar, Visscher

ENTM 253. Seminar in Insect Toxicology. (1) F Seminar, one hour. Prerequisite(s):ENTM 128 or consent of instructor. Selected topics in insect toxicology. Letter grades will be assigned to students presenting formal seminars: others will be graded Satisfactory (S) or No Credit (NC). Gill

ENTM 254. Seminar in Biological Control. (2) F,W Seminar, two hours. Prerequisite(s): BIOL 127/ ENTM 127, ENTM 129; or consent of instructor. Concepts, questions and hypotheses in biological control. Letter grades will be assigned to students presenting formal seminars; others will be graded Satisfactory (S) or No Credit (NC). Bellows

ENTM 255. Seminar in Medical and Veterinary Entomology. (2) F Seminar, two hours. Prerequisite(s): ENTM 126 or consent of instructor. Rigorous review and analysis of advanced topics in medical and veterinary entomology and related disciplines. Letter grades will be assigned to students presenting formal seminars; others will be graded Satisfactory (S) or No Credit (NC). Mulla, Mullens, Walton

ENTM 256. Seminar in Systematic Entomology. (2) S Seminar, two hours. Prerequisite(s): BIOL 112/ BPSC 112/ENTM 112 or consent of instructor. Selected topics in insect systematics. Letter grades will be assigned to students presenting formal seminars; others will be graded Satisfactory (S) or No Credit (NC). Heraty, Pinto

ENTM 258. Seminar in Insect Pest Management. (2) W Seminar, two hours. Prerequisite(s): consent of instructor. Selected topics in insect pest management. Letter grades will be assigned to students presenting formal seminars; by consent of the instructor, others will be graded Satisfactory (S) or No Credit (NC). Perring, Redak

ENTM 261. Colloquium in Recombinant DNA. (1) F,W,S Seminar, one hour. Prerequisite(s): graduate status or consent of instructor. Oral reports by visiting scholars, faculty and students on current research topics in recombinant DNA. Graded Satisfactory (S) or No Credit (NC). Course is repeatable. Cross-listed with BCH 261, BIOL 261, BPSC 261, and PLPA 261.

ENTM 271. Research Seminar in Management of Vegetable Crop Pests. (1) W Seminar, one hour. Prerequisite(s): consent of instructor. Seminar and critical discussion emphasizing current research and advances in management of vegetable crop pests. Graded Satisfactory (S) or No Credit (NC). Course may be repeated. Trumble

ENTM 272. Research Seminar in Insect Communication and Behavior. (1) F,W,S Seminar, one hour. Prerequisite(s): consent of instructor. Seminar and critical discussion emphasizing current research and advances in insect communication and behavior. Graded Satisfactory (S) or No Credit (NC). Course may be repeated. Cardé

ENTM 275. Research Seminar in Citrus Arthropod Pest Management. (1) W Seminar, one hour. Prerequisite(s): consent of instructor. Seminar and critical discussion emphasizing current research and advances in the management of arthropod pests of citrus and other subtropical tree crops. Graded Satisfactory (S) or No Credit (NC). Course is repeatable to a maximum of 4 units. Morse

ENTM 276. Research Seminar in Medical, Urban, and Veterinary Entomology. (1) F,S Seminar, one hour. Prerequisite(s): consent of instructor. Seminar and critical discussion emphasizing current research and advances in medical, urban, and veterinary entomology. Graded Satisfactory (S) or No Credit (NC). Course may be repeated. Mulla, Mullens, Rust, Walton

ENTM 277. Research Seminar in Insect Biochemistry and Toxicology. (1) F,W,S Seminar, one hour. Prerequisite(s): consent of instructor. Seminar and critical discussion emphasizing current research and advances in insect biochemistry and toxicology. Graded Satisfactory (S) or No Credit (NC). Course may be repeated. Gill

ENTM 281. Research Seminar in Molecular Entomology. (1) S, Even Years Seminar, one hour. Prerequisite(s): consent of instructor. Selected topics in molecular biology of insects and insect pathogens. Graded Satisfactory (S) or No Credit (NC). Atkinson

ENTM 289. Special Topics in Neuroscience. (2) F,W,S Seminar, two hours. Prerequisite(s): graduate standing or consent of instructor. An interdisciplinary seminar consisting of student presentations and discussion of selected topics in neuroscience. Content and instructor(s) vary each time course is offered. Letter grades will be assigned to students presenting formal seminars; others will be graded Satisfactory (S) or No Credit (NC). Course is repeatable. Cross-listed with BCH 289, BIOL 289, BMSC 289, CHEM 289, NRSC 289, and PSYC 289. Hatton in charge

ENTM 290. Directed Studies. (1-6) F,W,S Literature studies on special topics under direction of a member of the staff. Graded Satisfactory (S) or No Credit (NC). Course is repeatable.

ENTM 291. Individual Study in Coordinated Areas. (1-6) F,W,S Prerequisite(s): graduate standing. Faculty assisted programs of individual study for candidates who are preparing for examinations. The following rules apply: 1) Up to 6 units may be taken prior to award of the Master's degree, such units to be in addition to minimum unit requirements for the degree; 2) Up to 12 additional units may be taken prior to advancement to candidacy for the Ph.D.; 3) The course may be repeated within these limits; 4) Grading will be Satisfactory (S) or No Credit (NC).

ENTM 297. Directed Research (1-6) F,W,S Exploratory research toward the development of the dissertation problem or other research not specifically for thesis or dissertation. Graded Satisfactory (S) or No Credit (NC).

ENTM 299. Research for Thesis or Dissertation. (1-12) F,W,S Graded Satisfactory (S) or No Credit (NC). Course is repeatable.


PROFESSIONAL COURSES

ENTM 301. Teaching Entomology at the College Level. (1) F,W,S Seminar, one hour. Prerequisite(s): graduate standing in Entomology. A program of weekly meetings and individual formative evaluation required of new entomology Teaching Assistants. Covers instructional methods and classroom/section activities most suitable for teaching Entomology. Conducted by departmental faculty or the Teaching Assistant Development Program. Graded Satisfactory (S) or No Credit (NC). Course is repeatable.

ENTM 302. College Teaching Practicum. (1-4) F,W,S Practicum/consultation, three to twelve hours. Prerequisite(s): graduate standing and consent of instructor. Supervised teaching in college level classes under supervision of the course instructor. Graded Satisfactory (S) or No Credit (NC). Course is repeatable.